otseng wrote:Greetings Jose, it's been awhile since we've been head-to-head. Smile
Actually, I'm already stretched a bit too thin now....
And greetings to you, my friend. It has, indeed, been awhile. Actually, I'm already stretched a bit too thin now... but that's all just part of life's rich pageant, you know.
otseng wrote: Let's start off with this because it relates to how we're defining common descent. I provided a definition earlier: "all living organisms on Earth are descended from a single organism." For the purposes of this thread, this is how I'm referring to common descent.
but, the genetics and population biology for the common descent of all organisms from a common ancestor
follow the same rules as the common descent of all duck-like birds from a common ancestor of duck-like birds. There are, therefore, several issues here:
- do you accept that some species can have a common ancestor?
- do you accept the genetic mechanisms of inheritance of DNA from the parents, mutation of DNA, the dependence of phenotype on genotype, and the dependence of reproductive success on phenotype?
- what observations make common descent within certain animal groups acceptable, while the same mechanisms for "deeper" common descent between groups is unacceptable?
You see, genetically, it is impossible to separate deep-level common descent from recent, within-kind common descent. The data point to both levels being equally plausible, and identical in mechanism. I suspect that, like the Big Bang, the diversification of the first multicellular, soft-bodied animals is a target simply because there are fewer data. The precise route that evolution followed is not well known, so it's easy to say "okay,
show me." But, it's impossible to escape once one looks at biochemical, DNA-level relationships--the only other way to produce the data is to say "apparently god wanted to make everything look evolved."
otseng wrote:I'm not stating that there are no fossils prior to the Cambrian. Even in the chart above shows phyla prior to the Cambrian. ... Could you give references to exactly all the differing phyla that are found in the Precambrian?
That's a little like asking for the "current-animal name" of the common ancestor of cats and dogs. Since it was neither cat nor dog, it would not have a name as "cat" or "dog." Similarly, the precursor of two phyla would not fit into either, and would be called neither. Now, perhaps this reasoning indicates that the creationist argument that many phyla first appeared in the Cambrian represents nothing more than semantics. We couldn't
call them by extant phylum names until they kinda looked like 'em. There was lots of evolution before that, but the names were different.
In noodling about the literature, I've also learned that early paleontologists tried very hard to shoehorn Cambrian fossils into phyla that were known. Again, this is a matter of semantics. Does fossil XXX really belong to phylum chordata, or was it something else? What the hey--let's call it chordata anyway.
a blog on Darwin Central wrote: The phylum is the highest division except for kingdom, and these metazoans certainly belong in the animal kingdom. Since every living organism fits neatly into an extant phylum, there is a tendency to think that every extinct organism should fit neatly as well. --
http://www.darwincentral.org/blog/2006/ ... evolution/G. Budd wrote:One further aspect about these now extinct basal taxa is that they would have accumulated their own autapomorphies not possessed by the extant taxa. As a result, these basal fossil taxa are bound to differ from the extant clades: they will not be diagnosable as members of those clades; and they will show a confusing mixture of some but not all features of those clades, together with a set of features absent from them. It should be noted that this characteristic mix has been repeatedly noted in Cambrian fossils.
--The Cambrian fossil record and the origin of the phyla. Budd, G. Integrated Comprehensive Biology. 2003, 43, 157-165.
Because the fossils of soft-bodied things are rare and hard to find, and because the morphology of the earliest members of a phylum may be unrecognizably weird, many researchers have turned to DNA phylogeny to sort this out. This requires calibrating the molecular clock using branchpoints that are known from fossils of Cambrian age or younger, so there's a bit of extrapolation into the Precambrian. There are results such as this:
Wang et al wrote:In the past, molecular clocks have been used to estimate divergence times among animal phyla, but those time estimates have varied widely (1200-670 million years ago, Ma). In order to obtain time estimates that are more robust, we have analysed a larger number of genes for divergences among three well-represented animal phyla, and among plants, animals and fungi. The time estimate for the chordate-arthropod divergence, using 50 genes, is 993 +/- 46 Ma. Nematodes were found to have diverged from the lineage leading to arthropods and chordates at 1177 +/- 79 Ma. Phylogenetic analyses also show that a basal position of nematodes has strong support (p > 99%) and is not the result of rate biases. The three-way split (relationships unresolved) of plants, animals and fungi was estimated at 1576 +/- 88 Ma. By inference, the basal animal phyla (Porifera, Cnidaria, Ctenophora) diverged between about 1200-1500 Ma. This suggests that at least six animal phyla originated deep in the Precambrian, more than 400 million years earlier than their first appearance in the fossil record.
--
http://www.pubmedcentral.nih.gov/articl ... id=1689654
Proc Biol Sci. 1999 January 22; 266(1415): 163–171. Divergence time estimates for the early history of animal phyla and the origin of plants, animals and fungi. D Y Wang, S Kumar, and S B Hedges
otseng wrote:Jose wrote:It's unrealistic to expect (and biologically impossible anyway) that a new morphological feature will suddenly pop into existence in a single mutation. It will take multiple mutations over multiple generations to change a characteristic enough that we'd call it "new."
I think there are those who would disagree with this, but let's go with this. How can it empirically demonstrated that multiple mutations over multiple generations can generate something new?
Which part would they disagree with? Do they argue that there is hard data that show that new features
really do pop into existence with a single mutation? I bet not. Rather, I bet that they
claim that evolutionary theory posits this, and then wax poetic about what a dumb idea it is.
If a single mutation just can't do it, then it must require multiple mutations. Since we
know that a single new mutation cannot become fixed in a population without the passage of several generations, I think we're safe in saying that multiple mutations must require even more generations to become fixed. Since each mutation can do no more than change some existing trait, it must obviously require multiple changes before a trait can become different enough to call it "new."
Now, if people argue that there aren't enough mutations to have them accumulate over time, all they have to do is look at the DNA sequences of recently-diverged species. Take, for example,
Drosophila melanogaster and
Drosophila simulans. Look at the DNA sequences between genes. There are lots of differences. That's lots and lots of mutations.
otseng wrote:Jose wrote: I'd even question whether we should call things "new features" sometimes. Bird wings, bat wings, seal flippers, primate hands, and horse feet are all variations of the same thing: vertebrate limbs. And those are just variations of coelacanth fins, another variation of which is "ordinary" fish fins.
From my definition of common descent above, it is a requirement that new features be introduced in the evolutionary tree. If we go from a single cell organism to the variety of plants and animals on Earth, new features must have been introduced.
Indeed. But not in a single, magical leap in a single mutation in a single generation. The progenitor vertebrate limb had proximal bones and distal rays. In the lineage that became the ray-finned fish, the proximal bones were lost, and the rays expanded. In the lineage that became land animals, the rays were lost, and the bones expanded. This gives us fish fins and animal feet, one of which probably fits your definition of being "new." None of the individual changes were big enough to say "aha! from the last generation to this one, we've had the creation of a new feature." But after enough changes, we can look at the two lineages and say "gosh; these are different."
It's the same thing with "becoming multicellular." You start with cells being able to recognize each other and have what passes for sex (as in yeast). You move from there to cells being able to recognize each other and assemble into larger groups, as in stromatolites. From there you go to being able to adopt different "cell types" and play different roles in the loose assemblage of cells, as in sponges. From there, you go to more cell types and more complex roles...and here we are. No single change was very dramatic, but if we look at what we have now, compared to what we started with, whooooeee, lookit that!
otseng wrote: But, after some research, I've determined that the graph is unusable. And as I concluded in that post "there is no reason to doubt what the vast majority of biologists claim in that all the phyla appear during the Cambrian Explosion."
This is not a valid conclusion. The vast majority of biologists--at least, the vast majority of those I know--would go to UCMP to find out what the current thinking is. It doesn't matter what people used to think, except as an interesting study in how our understanding changes as more data are discovered.
otseng wrote: And there is no difference between 'micro-evolution' and 'macro-evolution' except that micro is within a few generations, and 'macro' is over hundreds of generations.
Looking at the Cambrian fossils, we do see examples of microevolution among the trilobites. There is a great diversity among the types of trilobites. However, we see no evidence of trilobites evolving into something else. Or any other animal evolving into something else. If microevolution spans hundreds or even thousands of generations, we should see evidence of a smooth and gradual evolution between various animals. However, fossils that we see are distinct and quite different from the others. I give some of the fossils found in the Cambrian here.
"Microevolution" is the change in allele frequencies; "macroevolution" is a longer-term change in morphology, including speciation and higher-level diversification. The trilobites certainly diversified into multiple genera and even families before they succumbed to the Permian extinction. They might well have given rise to other types of things had they not become extinct.
Again, it's no more compelling to say "Cambrian fossils are all distinct types" than to say "today's animals are all distinct," or to say "Ediacaran fossils are all distinct." The only way to justify claiming that organisms appeared suddenly with no prior evolution is if there are no prior fossils. That's what created the myth of the Cambrian Explosion--the fact that it took a long time to find Precambrian fossils, so it kinda looked like, maybe, there were no prior animals that could have been ancestors. Once we know there are fossils, and know there were prior life forms, we can no longer argue with any shred of validity that the Cambrian critters had no ancestors. Rather, the question becomes, "which of the fossils could possibly represent the ancestors of the Cambrian critters? Or, which could be relatives of the direct ancestors?" The puzzle now is to figure out what an animal might be like that had only one or two (ancestral) Hox genes, rather than multiple clusters of Hox genes in a large gene family.